Background and activities


I am a marine behavioural ecologist with a particular interest in benthic marine invertebrates. My primary research goals are aimed at understanding how environmental conditions influence marine animal behaviour, determining the proximate mechanisms mediating such influences, and elucidating the functional ramifications of environmental effects on behaviour. The environmental conditions I'm interested in can be biotic (e.g. threat of predation, conspecific associations, parasitism) or abiotic (environmental conditions), and can be natural (e.g. food availability, seasonal changes in environmental parameters, natural predators) or anthropogenic (e.g. ocean acidification, ocean warming, deoxygenation, anthropogenically-introduced species). The questions I study span the fundamental-applied research continuum, and are often both exploratory and applied.


I am also highly interested in experimental pedagogy and open access scholarly publishing. 




Clements JC, Comeau LA. 2019. Behavioural defenses of shellfish prey under ocean acidification. J. Shellfish Res. In press.

Clements JC, Comeau LA. 2019. Use of an emerging technology to detect ocean acidification effects on shellfish behaviour. J. Shellfish Res. In press.

Carroll J, Clements JC. 2019. Scaredy-oysters: in situ documentation of an oyster behavioral response to predators. Southeast. Nat. In press.

Kong H, Clements JC, Dupont S, Wang T, Huang, X, Shang Y, Liu L, Huang W, Hu M, Wang Y. 2019. Seawater acidification and temperature modulate anti-predator defenses in two co-existing Mytilus species. Mar. Poll. Bull. 145:118-125.

Kong H, Jiang X, Clements JC, Wang T, Huang X, Shang Y, Chen J, Hu M, Wang Y. 2019. Transgenerational effects of short-term exposure to acidification and hypoxia on early developmental traits of the mussel Mytilus edulis. Mar. Environ. Res. 145:73-80.

Klompmaker AA, Kelley PH, Chattopadhyay D, Clements JC, Huntley JW, Kowalewski M. 2019. Predation in the marine fossil record: Studies, data, recognition, environmental factors, and behavior. Earth Sci. Rev. 194:472-520.

Poirier LA, Clements JC, Davidson JDP, Miron G, Davidson J, Comeau LA. 2019. Sink before you settle: settlement behaviour of Eastern oyster (Crassostrea virginica) larvae on artificial spat collectors and natural substrate. Aquacult. Rep. 13:100181. 

Clements JC, Comeau LA. 2019. Nitrogen removal potential of shellfish aquaculture harvests in eastern Canada: A comparison of culture methods. Aquacult. Rep. 13:100183.



Clements JC, Hicks C, Tremblay R, Comeau LA. 2018. Elevated seawater temperature, not CO2, affects energetic reserves, body condition, and byssal attachment strength in adult mussels (Mytilus edulis) under food limitation. Con. Physiol. 6:cox078.

Clements JC, Darrow ES. 2018. Eating in an acidifying ocean: a quantitative review of elevated CO2 effects on the feeding rates of calcifying marine invertebrates. Hydrobiologia. 820:1-21.

Clements JC, Coffin MRS, Lavaud R, Guyondet T, Comeau LA. 2018. Ocean acidification and molluscan shell taphonomy: can elevated seawater pCO2 influence taphonomy in a naticid predator–prey system? Paleogeogr, Paleoclimatol, Paleoecol. 507:145-154.

Clements JC, Hunt HL. 2018. Testing for sediment acidification effects on within-season variability in juvenile soft-shell clam (Mya arenaria) abundance on the northern shore of the Bay of Fundy. Estuar. Coasts 41:471-183.

Clements JC, Bourque D, McLaughlin J, Stephenson M, Comeau L. 2018. Wanted dead or alive: Polydora websteri recruit on both live oysters and empty shells of the eastern oyster, Crassostrea virginica. J. Fish. Diseases. 41:855-858.

Clements JC, Daigle RM, Froehlich HE. 2018. Predator in the pool? A quantitative evaluation of non-indexed open access journals in aquaculture research. Front. Mar. Sci. 5:106.

Clements JC, Davidson JDP, Gunn McQuillan J, Comeau LA. 2018. Increased mortality of harvested eastern oysters (Crassostrea virginica) is associated with air exposure and temperature during a spring fishery in Atlantic Canada. Fisheries Res. 206:27-34.

Clements JC, Comeau LA, Carver CE, Mayrand E, Plante S, Mallet A. 2018. Short-term exposure to elevated pCO2 does not affect the valve gaping response of adult eastern oysters, Crassostrea virginica, to acute heat shock under an ad libitum feeding regime. J. Exp. Mar. Biol. Ecol. 506:9-17.



Clements JC, Bishop MM, Hunt HL. 2017. Elevated temperature has adverse effects on GABA-mediated adaptive behaviour to sediment acidification in a wide-ranging marine bivalve. Mar. Biol. 164:56.

Clements JC, Hunt HL. 2017. Effects of CO2-driven sediment acidification on infaunal marine bivalves: a synthesis. Mar. Poll. Bull. 117:6-16.

Clements JC, Bourque D, McLaughlin J, Stephenson M, Comeau L. 2017. Extreme ocean acidification reduces the susceptibility of eastern oyster shells to a polydorid parasite. J. Fish Diseases. 40:1573-1585.

Clements JC, Bourque D, McLaughlin J, Stephenson M, Comeau L. 2017. Siltation increases the susceptibility of surface-cultured eastern oysters (Crassostrea virginica) to parasitism by the mudworm Polydora websteri. Aquaculture Res. 48 :4707-4717.

Ferguson LF, Davidson JDP, Landry T, Clements JC, Therriault TW. 2017. Didemnum vexillum: Invasion potential via harvesting and processing of the Pacific oyster (Crassostrea gigas) in British Columbia, Canada. Manag. Biol. Invasions 8:553-558.

Clements JC, McCorquodale DB, Doucet BA, Ogden JB. 2017. The Dusky Cockroach in the Canadian Maritimes: establishment, persistence, and ecology. J. Acad. Entomol. Soc. 13:21-27.

Clements JC. 2017. Open access articles receive more citations in hybrid marine ecology journals. FACETS 2:1-14.



Clements JC, Chopin T. 2016. Ocean acidification and marine aquaculture in North America: potential impacts and mitigation strategies. Rev. Aquaculture 9:326-341.

Clements JC, Woodard KW, Hunt HL. 2016. Porewater acidification alters the burrowing behaviour and post-settlement dispersal of juvenile soft-shell clams (Mya arenaria). J. Exp. Mar. Biol. Ecol. 447:103-111.



Clements JC. 2015. Using Facebook to enhance independent student engagement: a case study with first-year undergraduates. Higher Educ. Stud. 5:131-146.

Clements JC, Hunt HL. 2015. Marine animal behavior in a high-CO2 ocean. Mar. Ecol. Prog. Ser. 536: 259-279.



Clements JC, Hunt HL. 2014. Influence of sediment acidification and water flow on sediment acceptance and dispersal of juvenile soft-shell clams (Mya arenaria L.). J. Exp. Mar. Biol. Ecol. 453:62-69.

Ghasemi F, Clements JC, Taheri M, Rostami A. 2014. Changes in quantitative distribution of Caspian Sea polychaetes: prolific fauna formed by non-indigenous species. J. Great Lakes Res. 40:692-698.

Clements JC, Rawlings TA. 2014. Ontogenetic shifts in the predatory habits of the northern moonsnail (Lunatia heros) on the northwestern Atlantic coast. J. Shellfish Res. 33:755-768.



Clements JC, Doucet DA, McCorquodale DB. 2013. Establishment of a European cockroach, Ectobius lapponicus (L.) (Dictyoptera: Blattellidae), in the Maritime Provinces of eastern Canada. J. Acad. Entomol. Soc. 9:4-7.

Clements JC, Ellsworth-Power M, Rawlings T. 2013. Diet breadth of the northern moonsnail (Lunatia heros) on the northwestern Atlantic coast. Am. Mal. Bull. 31:331-336.



Clements JC. 2012. First record of the Fiery Skipper, Hylephila phyleus Drury (Lepidoptera: Hesperiidae) from New Brunswick, Canada. J. Acad. Entomol. Soc. 8:55-60.

Scientific, academic and artistic work