2026

• Hansson Frank, Tor; Araya Ajoy, Yimen Gerardo; Wright, Jonathan; Sæther, Bernt-Erik; Jensen, Henrik. (2026) Density-dependent evolutionary demography in a small passerine – from the metapopulation level to life-history traits. Norges teknisk-naturvitenskapelige universitet
  Doctoral thesis
• Réale, Denis; Allegue, Hassen; Araya Ajoy, Yimen Gerardo; Dochtermann, Ned A.; Nakagawa, Shinichi; Pick, Joel L.; Schielzeth, Holger; Westneat, David F.; Dingemanse, Niels J.. (2026) Avoiding misleading estimates of among-individual variance caused by non-random sampling of individuals in a changeable environment. Methods in Ecology and Evolution
  Academic article
• Hansson Frank, Tor; Ranke, Peter Sjolte; Burnett, Hamish Andrew; Acker, Paul; Ringsby, Thor Harald; Wright, Jonathan; Jensen, Henrik; Sæther, Bernt-Erik; Araya Ajoy, Yimen Gerardo. (2026) Spatial heterogeneity in density dependence across life-history stages drives the dynamics of a bird metapopulation. Ecology
  Academic article

2025

• Class, Barbara; Araya Ajoy, Yimen Gerardo; Dingemanse, Niels J.; Dochtermann, Ned A.; Martin, Jordan S.; Moiron, Maria; Westneat, David F.. (2025) Disentangling nonrandom assortment, indirect effects, and joint plasticity as causes of phenotypic (dis)similarity between social partners. Journal of Evolutionary Biology
  Academic article
• Martin, Jordan S.; Westneat, David F.; Wilson, Alastair J.; Dingemanse, Niels J.; Araya Ajoy, Yimen Gerardo. (2025) Frequency dependence favours social plasticity and facilitates socio-eco-evolutionary feedback in fluctuating environments. Functional Ecology
  Academic article
• Martin, Jordan S.; Araya Ajoy, Yimen Gerardo; Dingemanse, Niels J.; Wilson, Alastair J.; Westneat, David F.. (2025) Estimating (Non)Linear Selection on Reaction Norms: A General Framework for Labile Traits. Ecology and Evolution
  Academic article
• Pélabon, Christophe; Agudelo-Cantero, Gustavo Adolfo; Araya Ajoy, Yimen Gerardo; Bolstad, Geir Hysing; Cheng, Changde; Galis, Frietson; Guillaume, Frederic; Haaland, Thomas Ray; Hallgrímsson, Benedikt; Hansen, Thomas Fredrik. (2025) Evolvability: progress and key questions. BioScience
  Academic article
• Araya Ajoy, Yimen Gerardo; Murray, Myranda; Engen, Steinar; Sæther, Bernt-Erik; Wright, Jonathan. (2025) Decomposing Frequency Dependence on Selection and Population Growth: Implications for Carrying Capacity and Maladaptation. Ecology Letters
  Academic article
• Goedert, Debora; Ringsby, Thor Harald; Jensen, Henrik; Kvalnes, Thomas; Araya Ajoy, Yimen Gerardo; Skomedal, Silje. (2025) Relationships between weather, arthropod biomass, and natal dispersal in house sparrows (Passer domesticus) in northern Norway. Norges teknisk-naturvitenskapelige universitet
  Master thesis
• Araya Ajoy, Yimen Gerardo; Hansson Frank, Tor; Burnett, Hamish Andrew; Søraker, Jørgen Skavdal; Ranke, Peter Sjolte; Goedert, Debora; Ringsby, Thor Harald; Jensen, Henrik; Sæther, Bernt-Erik. (2025) Assessing the 'Small Population' Paradigm: The Effects of Stochasticity on Evolutionary Change and Population Growth in a Bird Metapopulation. Ecology Letters
  Commentary
• Saatoglu, Fikriye Dilan; Niskanen, Alina Katariina; Froy, Hannah; Ranke, Peter Sjolte; Goedert, Debora; Reid, Jane Margaret; Ringsby, Thor Harald; Sæther, Bernt-Erik; Araya Ajoy, Yimen Gerardo; Jensen, Henrik. (2025) Metapopulation-level analyses reveal positive fitness consequences of immigration in a small bird. Journal of Animal Ecology
  Academic article

2024

• Sandoval, Luis; Graham, Brendan; Sosa-López, J. Roberto; Laverde-R., Oscar; Araya Ajoy, Yimen Gerardo. (2024) THE STUDY OF BIRD VOCALIZATIONS IN NEOTROPICAL HABITATS: CURRENT KNOWLEDGE AND FUTURE STEPS. Ornitologia Neotropical
  Academic article
• Ranke, Peter Sjolte; Pepke, Michael Le; Søraker, Jørgen Skavdal; David, Gabriel; Araya Ajoy, Yimen Gerardo; Wright, Jonathan; Nafstad, Ådne Messel; Rønning, Bernt; Pärn, Henrik; Ringsby, Thor Harald. (2024) Long-distance dispersal in the short-distance dispersing house sparrow (Passer domesticus). Ecology and Evolution
  Academic article
• Murray, Myranda; Wright, Jonathan; Araya Ajoy, Yimen Gerardo. (2024) Evolutionary rescue from climate change: male indirect genetic effects on lay-dates and their consequences for population persistence. Evolution Letters
  Academic article
• Chaverri, Gloriana; Sagot, Maria; Stynoski, Jennifer L.; Araya-Salas, Marcelo; Araya Ajoy, Yimen Gerardo; Nagy, Martina; Knörnschild, Mirjam; Chaves-Ramírez, Silvia; Rose, Nicole; Sánchez-Chavarría, Mariela. (2024) Calling to the collective: Contact calling rates within groups of disc-winged bats do not vary by kinship or association. Philosophical Transactions of the Royal Society of London. Biological Sciences
  Academic article
• Gamelon, Marlene; Araya Ajoy, Yimen Gerardo; Sæther, Bernt-Erik. (2024) The concept of critical age group for density dependence: Bridging the gap between demographers, evolutionary biologists and behavioural ecologists. Philosophical Transactions of the Royal Society of London. Biological Sciences
  Academic article

2023

• Pick, Joel; Kasper, Claudia; Allegue, Hassen; Dingemanse, Niels J.; Dochtermann, Ned A.; Laskowski, Kate L.; Lima, Marcos R.; Schielzeth, Holger; Westneat, David F.; Wright, Jonathan. (2023) Describing posterior distributions of variance components: Problems and the use of null distributions to aid interpretation. Methods in Ecology and Evolution
  Academic article
• Groot, Corné de; Wijnhorst, Rori E.; Ratz, Tom; Murray, Myranda Elizabeth; Araya Ajoy, Yimen Gerardo; Wright, Jonathan; Dingemanse, Niels J.. (2023) The importance of distinguishing individual differences in ‘social impact’ versus ‘social responsiveness’ when quantifying indirect genetic effects on the evolution of social plasticity. Neuroscience and Biobehavioral Reviews
  Academic literature review
• Peignier, Mélissa; Araya Ajoy, Yimen Gerardo; Ringler, Max; Ringler, Eva. (2023) Personality traits differentially affect components of reproductive success in a Neotropical poison frog. Proceedings of the Royal Society of London. Biological Sciences
  Academic article
• Pepke, Michael Le; Kvalnes, Thomas; Wright, Jonathan; Araya Ajoy, Yimen Gerardo; Ranke, Peter Sjolte; Boner, Winnie; Monaghan, Pat; Sæther, Bernt-Erik; Jensen, Henrik; Ringsby, Thor Harald. (2023) Longitudinal telomere dynamics within natural lifespans of a wild bird. Scientific Reports
  Academic article
• Araya Ajoy, Yimen Gerardo; Dingemanse, Niels J.; Westneat, David F.; Wright, Jonathan. (2023) The evolutionary ecology of variation in labile traits: selection on its among- and within-individual components. Evolution
  Academic article
• Munar-Delgado, Gabriel; Araya Ajoy, Yimen Gerardo; Edelaar, Pim. (2023) Estimation of additive genetic variance when there are gene–environment correlations: Pitfalls, solutions and unexplored questions. Methods in Ecology and Evolution
  Academic article

2022

• Peignier, Mélissa; Araya-Ajoy, Yimen; Bégué, Lauriane; Chaloupka, Sarah; Dellefont, Katharina; Leeb, Christoph; Walsh, Patrick; Ringler, Max; Ringler, Eva. (2022) Exploring links between personality traits and their social and non-social environments in wild poison frogs. Behavioral Ecology and Sociobiology
  Academic article
• Chaloupka, Sarah; Peignier, Mélissa; Stückler, Susanne; Araya-Ajoy, Yimen; Walsh, Patrick; Ringler, Max; Ringler, Eva. (2022) Repeatable Territorial Aggression in a Neotropical Poison Frog. Frontiers in Ecology and Evolution
  Academic article
• Pepke, Michael Le; Kvalnes, Thomas; Ranke, Peter Sjolte; Araya-Ajoy, Yimen; Wright, Jonathan; Sæther, Bernt-Erik; Jensen, Henrik; Ringsby, Thor Harald. (2022) Causes and consequences of variation in early-life telomere length in a bird metapopulation. Ecology and Evolution
  Academic article

2021

• Saatoglu, Fikriye Dilan; Niskanen, Alina Katariina; Kuismin, Markku; Ranke, Peter Sjolte; Hagen, Ingerid Julie; Araya-Ajoy, Yimen; Kvalnes, Thomas; Pärn, Henrik; Rønning, Bernt; Ringsby, Thor Harald. (2021) Dispersal in a house sparrow metapopulation: An integrative case study of genetic assignment calibrated with ecological data and pedigree information. Molecular Ecology
  Academic article
• Muller, Amandine Sophie; Jensen, Henrik; Araya-Ajoy, Yimen; Husby, Arild; Kvalnes, Thomas; Lundregan, Sarah; Niskanen, Alina Katariina; Ringsby, Thor Harald; Wright, Jonathan. (2021) Effects of inbreeding on lay date in house sparrows (Passer domesticus). Norges teknisk-naturvitenskapelige universitet
  Master thesis
• Araya-Ajoy, Yimen; Niskanen, Alina Katariina; Froy, Hannah; Ranke, Peter Sjolte; Kvalnes, Thomas; Rønning, Bernt; Pepke, Michael Le; Jensen, Henrik; Ringsby, Thor Harald; Sæther, Bernt-Erik. (2021) Variation in generation time reveals density regulation as an important driver of pace of life in a bird metapopulation. Ecology Letters
  Academic article
• Dingemanse, Niels J.; Araya-Ajoy, Yimen; Westneat, David F.. (2021) Most published selection gradients are underestimated: Why this is and how to fix it. Evolution
  Academic article
• Ranke, Peter Sjolte; Araya-Ajoy, Yimen; Ringsby, Thor Harald; Pärn, Henrik; Rønning, Bernt; Jensen, Henrik; Wright, Jonathan; Sæther, Bernt-Erik. (2021) Spatial structure and dispersal dynamics in a house sparrow metapopulation. Journal of Animal Ecology
  Academic article

2020

• Lundin, Thorbjørn; Wright, Jonathan; Araya-Ajoy, Yimen. (2020) Network analysis of genetic relatedness using population dynamic simulations. Norges teknisk-naturvitenskapelige universitet
  Master thesis
• O'Shea, Myranda; Wright, Jonathan; Araya-Ajoy, Yimen. (2020) The Role of Indirect Genetic Effects in Evolutionary Rescue. Norges teknisk-naturvitenskapelige universitet
  Master thesis
• Dingemanse, Niels; Moiron, Maria; Araya-Ajoy, Yimen; Mouchet, Alexia; Abbey-Lee, Robin. (2020) Individual variation in age-dependent reproduction: Fast explorers live fast but senesce young?. Journal of Animal Ecology
  Academic article
• Schielzeth, Holger; Dingemanse, Niels J.; Nakagawa, Shinichi; Westneat, David F.; Allegue, Hassen; Teplitsky, Céline; Réale, Denis; Dochtermann, Ned A.; Garamszegi, László Zsolt; Araya-Ajoy, Yimen. (2020) Robustness of linear mixed-effects models to violations of distributional assumptions. Methods in Ecology and Evolution
  Academic article
• Moiron, Maria; Araya-Ajoy, Yimen; Teplitsky, Céline; Bouwhuis, Sandra; Charmantier, Anne. (2020) Understanding the social dynamics of breeding phenology: indirect genetic effects and assortative mating in a long-distance migrant. The American Naturalist
  Academic article
• Chaverri, Gloriana; Araya-Ajoy, Yimen; Sagot, Maria. (2020) Contact calling in context: intra- and intergroup variation in vocalization rates depend on a call’s function. Behavioral Ecology and Sociobiology
  Academic article
• Westneat, David F.; Araya-Ajoy, Yimen; Allegue, Hassen; Class, Barbara; Dingemanse, Niels; Dochtermann, Ned A.; Garamszegi, László Zsolt; Martin, Julien G.A.; Nakagawa, Shinichi; Réale, Denis. (2020) Collision between biological process and statistical analysis revealed by mean centring. Journal of Animal Ecology
  Academic article
• Juárez, Roselvy; Araya-Ajoy, Yimen; Barrantes, Gilbert; Sandoval, Luis. (2020) House Wrens Troglodytes aedon reduce repertoire size and change song element frequencies in response to anthropogenic noise. IBIS
  Academic article
• Araya-Ajoy, Yimen; Westneat, David F.; Wright, Jonathan. (2020) Pathways to social evolution and their eco-evolutionary feedbacks. Evolution
  Academic article
• Niskanen, Alina Katariina; Billing, Anna Maria; Holand, Håkon; Hagen, Ingerid Julie; Araya-Ajoy, Yimen; Husby, Arild; Rønning, Bernt; Myhre, Ane Marlene; Ranke, Peter Sjolte; Kvalnes, Thomas. (2020) Consistent scaling of inbreeding depression in space and time in a house sparrow metapopulation. Proceedings of the National Academy of Sciences of the United States of America (PNAS)
  Academic article
• Engen, Steinar; Wright, Jonathan; Araya-Ajoy, Yimen; Sæther, Bernt-Erik. (2020) Phenotypic evolution in stochastic environments: The contribution of frequency- and density-dependent selection. Evolution
  Academic article
• Jensen, Henrik; Araya-Ajoy, Yimen; Lundregan, Sarah; Samu, Attila. (2020) Spatio-temporal variation in phenotypic selection in a metapopulation of house sparrows (Passer domesticus). Norges teknisk-naturvitenskapelige universitet
  Master thesis

2019

• Moiron, Maria; Araya-Ajoy, Yimen; Mathot, Kimberley J; Mouchet, Alexia; Dingemanse, Niels J.. (2019) Functional relations between body mass and risk-taking behavior in wild great tits. Behavioral Ecology
  Academic article
• Mutzel, Ariane; Olsen, Anne-Lise; Mathot, Kimberley J; Araya-Ajoy, Yimen; Nicolaus, Marion; Wijmenga, Jan J; Wright, Jonathan; Kempenaers, Bart; Dingemanse, Niels J.. (2019) Effects of manipulated levels of predation threat on parental provisioning and nestling begging. Behavioral Ecology
  Academic article
• Wright, Jonathan; Bolstad, Geir Hysing; Araya-Ajoy, Yimen; Dingemanse, Niels J.. (2019) Life-history evolution under fluctuating density-dependent selection and the adaptive alignment of pace-of-life syndromes. Biological Reviews
  Academic article
• Araya-Ajoy, Yimen; Ranke, Peter Sjolte; Kvalnes, Thomas; Rønning, Bernt; Holand, Håkon; Myhre, Ane Marlene; Pärn, Henrik; Jensen, Henrik; Ringsby, Thor Harald; Sæther, Bernt-Erik. (2019) Characterizing morphological (co)variation using structural equation models: Body size, allometric relationships and evolvability in a house sparrow metapopulation. Evolution
  Academic article
• Fried-Petersen, Hannah; Araya-Ajoy, Yimen; Futter, Martyn N.; Angeler, David G.. (2019) Drivers of long-term invertebrate community stability in changing Swedish lakes. Global Change Biology
  Academic article
• Jolles, Jolle W; Briggs, Helen D; Araya-Ajoy, Yimen; Boogert, Neeltje. (2019) Personality, plasticity and predictability in sticklebacks: bold fish are less plastic and more predictable than shy fish. Animal Behaviour
  Academic article

2018

• Abbey-Lee, Robin N; Araya-Ajoy, Yimen; Mouchet, Alexia; Moiron, Maria; Stuber, Erica F; Kempenaers, Bart; Dingemanse, Niels J.. (2018) Does perceived predation risk affect patterns of extra‐pair paternity? A field experiment in a passerine bird. Functional Ecology
  Academic article
• Araya-Ajoy, Yimen; Bolstad, Geir Hysing; Brommer, Jon; Careau, Vincent; Dingemanse, Niels J.; Wright, Jonathan. (2018) Demographic measures of an individual's "pace of life": fecundity rate, lifespan, generation time, or a composite variable?. Behavioral Ecology and Sociobiology
  Academic article
• Sonnweber, Ruth; Araya-Ajoy, Yimen; Behringer, Verena; Deschner, Tobias; Tkaczynski, Patrick; Fedurek, Pawel; Preis, Anna; Samuni, Liran; Zommers, Zinta; Gomes, Cristina. (2018) Circadian rhythms of urinary cortisol levels vary between individuals in wild male chimpanzees: A reaction norm approach. Frontiers in Ecology and Evolution
  Academic article

2017

• Class, Barbara; Dingemanse, Niels J.; Araya-Ajoy, Yimen; Brommer, Jon E.. (2017) A statistical methodology for estimating assortative mating for phenotypic traits that are labile or measured with error. Methods in Ecology and Evolution
  Academic article
• Serrano-Davies, Eva; Araya-Ajoy, Yimen; Dingemanse, Niels J.; Sanz, Juan Jose. (2017) Personality-related differences in response to habitat in Mediterranean blue tits. Ethology
  Academic article
• Mathot, Kimberley J; Olsen, Anne-Lise; Mutzel, Ariane; Araya-Ajoy, Yimen; Nicolaus, Marion; Westneat, David F.; Wright, Jonathan; Kempenaers, Bart; Dingemanse, Niels J.. (2017) Provisioning tactics of great tits (Parus major) in response to long-term brood size manipulations differ across years. Behavioral Ecology
  Academic article
• Allegue, Hassen; Araya-Ajoy, Yimen; Dingemanse, Niels J.; Dochtermann, Ned A.; Garamszegi, László Zsolt; Nakagawa, Shinichi; Réale, Denis; Schielzeth, Holger; Westneat, David F.. (2017) Statistical Quantification of Individual Differences (SQuID): an educational and statistical tool for understanding multilevel phenotypic data in linear mixed models. Methods in Ecology and Evolution
  Academic article
• Araya-Ajoy, Yimen; Dingemanse, Niels J.. (2017) Repeatability, heritability, and age-dependence of seasonal plasticity in aggressiveness in a wild passerine bird. Journal of Animal Ecology
  Academic article

2016

• Araya-Ajoy, Yimen; Kuhn, Sylvia; Mathot, Kimberley J; Mouchet, Alexia; Mutzel, Ariane; Nicolaus, Marion; Wijmenga, Jan J; Kempenaers, Bart; Dingemanse, Niels J.. (2016) Sources of (co)variation in alternative siring routes available to male great tits (Parus major). Evolution
  Academic article