Background and activities

Dr. James D. M. Speed

My research involves a range of taxa from vascular plants and bryophytes to invertebrates, birds and large mammalian herbivores. I work at a range of scales, from the individual to the ecosystem, with short-term experiments to palaeoecology, and from local scale patterns to regional scale processes. I have worked in environments from the high-arctic and temperate-alpine regions to boreal forests and African savannahs. A core interest is the impact of vertebrate herbivores and land-use on plant populations and communities. My work also focuses on the ability of herbivores to affect ecosystem and vegetation states and the how the impact of herbivory on ecosystems feeds back to herbivore populations.

I am a fellow on the NTNU Outstanding Academic Fellows programme and a holder of a Research Council of Norway Young Research Talents project.

I am a member of the editorial board for the journals Ecology and Ecological Monographs

Current Research

Disentangling the impacts of herbivory and climate on ecological dynamics: The DISENTAGLE project addresses how climate and herbivory together affect ecosystem processes across spatial scales. The project investigates interactions between herbivory and climate in northern ecosystems by examining (1) The growth of woody plants and community composition at the site scale. (2) The ability of herbivores to determine plant species distributions at a regional scale and (3) The functional and phylogenetic ecology of herbivores across the world's arctic and boreal ecosystems.

CHARTER: Drivers and feedbacks of changes in Arctic terrestrial biodiversity.

ForestPotential Changing Forest Area and Forest Productivity – Climatic and Human Causes, Effects, Monitoring Options, and Climate Mitigation Potential

SustHerb Project The SustHerb project is working towards sustainable management of moose, red deer and their food resources.  I am involved in analysing vegetation and tree response to browsing herbivores.

Previous projects

AfricanBioServices: Investigating how productivity, nutrient cycling and carbon storage vary with livestock and wild herbivore grazing along a climatic gradient in the Serengeti-Mara ecosystem

MIRACLE project: Mires and climate: towards enhancing functional resilience of fen peatlands

Sheep Grazing Project Quantification of the impact of grazing on the distribution and growth of mountain birch above the forest line. 

Managing ecosystem services in low alpine cultural landscapes through livestock grazing Moving on from this project we  are investigating how sheep grazing impacts on alpine ecosystem services through impacts on the treeline.

Systematic review of impacts of reindeer grazing. I was part of a team carrying out a systematic review of the impacts of reindeer herbivory on mountain and arctic vegetation 

DYLAN Project Homepage I also worked within the DYLAN project where I am synthesising the impact of land use and natural drivers of landscape state across protected areas in Norway and the UK, with a long-term perspective. 

Former Postdocs and Researchers

Stefaniya Kamenova (Postdoc 2021- ) Metabarcoding of reindeer diet in relation to changes in Arctic terrestrial biodiversity

Tanja K. Petersen (Researcher 2021-2022) Alpine ecology, functional biogeography, natural history collections

Stuart W. Smith (Postdoc 2015-2019) Investigating how wildlife and livestock affect vegetation and ecosystem processes in the Serengeti-Mara region in Tanzania and Kenya

Kenny Helsen (Postdoc 2015-2017) Assessing the impact of invasive plant species on ecosystem services and associated ecosystem functions using a functional effect trait framework.

PhD Students

Mathilde Defourneaux (PhD student 2021.2024, Agricultural University of Iceland, main supervisor Isabel Barrio). Herbivores in the tundra: Linking diversity and function.

Ann Evankow (PhD student 2020-2023 University of Oslo) Integrative taxonomy and spatial phylogenetics of the Psoraceae

Katy Ivison (PhD student 2019-2022, Durham University, main supervisor is Wayne Dawson) Enemies of the invaders: How will natural enemies affect the risk of non-native plant invasions in a warmer Arctic?

Damaris M. Matten (PhD student 2019-2022). Ecology and phylogenetics

Marte Fandrem (PhD student 2018-2022) Peat mosses and ecosystem services

Former PhDs

Malene Ø. Nygård (PhD student 2017-2021, main supervisor is Mika Bendiksby) Plant conservation genomics

Katariina Vuorinen (PhD student 2017-2021) When do ungulates override the climate? Defining the interplay of two key drivers of northern vegetation dynamics.

Tanja K. Petersen (PhD student 2017-2021, main supervisor is Gunnar Austrheim) Biodiversity dynamics in urban areas under changing land-uses

Anders L. Kolstad (PhD student 2015-2019) Ungulate-mediated effects on ecosystem functions and services in boreal production forests

Master Students

Anna Sobocinksi (Master student (2020-2022) Mapping carnivore community structure in Norway across 100 years

Emma Petterson (Master student 2020-2021) Functional diversitsy of Arctic carnviores

Sonia Pujara (Master student 2020-2022) How do warming and herbivory affect the phenology of non-native plant species?

Rachel Guindon (Master student Université Laval, Québec, 2018-2020, main supervisor is Jean-Pierre Tremblay) Assessment of the resources used by muskoxen (Ovibos moschatus) in Nunavik and development of a community monitoring protocol.

John Maxwell Salisbury (Master student 2019-2021) How do moose influence climate? The effect of moose browsing on surface albedo in Norwegian successional boreal forest.

Former master students

Peter Frank (Master student 2018-2020) Climate and herbivory as drivers of woody plant growth in Alaksa

Maria Alejandra Aguilar Tomasini (Master student 2018-2020) The phylogenetic diversity of Mexican mammals

Hans Ivar Hortman (Master Student 2018-2020) Climate and reindeer herbivory as drivers of woody plant growth in Finnmark.

Kjersti Rønhovde Styve (Master student 2019-2020) Phylogenetic Diversity Patterns across Trophic Levels in Norway.

Ingrid Bekken Snøan (Master student 2018-2019) Use of Lidar data in assessing effect of moose browsing on boreal forests

Jesus Adrian Chimal Ballesteros (Master student 2017-2019) Mapping the functional and phylogenetic diversity of boreal and arctic herbivores

Vilde Lytskjold Haukenes (Master student 2017-2019) Impacts of climate and land-use change on carbon storage in the Serengeti-Mara ecosystem

Sarah Adom Yawsom (Master student 2017-2019) Integrating ecological informatics and ethnobotany to map ecosystem services in Ghana

Ida Marielle Mienna (Master student 2016-2018) Spatial phylogenetics of the Norwegian flora.

Victoria Berger (Master student 2016-2018) Successional dynamics in boreal forests: what is the impact of moose browsing, and which pathways are preferred by landowners?

Winta Gebreyohanis (Master student 2015-2017) The effects of browsing and climate on above- ground carbon stocks in boreal forests.

Joana Awuah Adofo (Master student 2015-2017) The impact of bushfire on plant functional groups and carbon storage in the savanna of northern Ghana.

Philbert M. Ngoti (Master student 2015-2017) Feeding ecology Of Eastern Black Rhinoceros (Diceros Bicornis Michaeli) in southern Serengeti National Park, Tanzania.

Ina Åsnes Skjellbred (Master student 2016-2017) Spatial patterns in the phylogenetic diversity of Arctic herbivores.

Toril Nes (Master student 2013 – 2015). The Effects of Moose (Alces alces)Browsing on Boreal Tree Species in Norway and Quebec

Kristin Wangen (Master student 2013 – 2015). Understanding the Ecology of Three Mixed Northern Hepatic Mat Species at Regional Scale through Species Distribution Modelling, and Local Scale through Growth Measurements and Micro-Climatic Assessment.

Marie Maurset (Master student 2013 – 2015). Long-term Effects of Shifts in Grazing Pressure on Alpine Plant Species along an Elevational Gradient.

Else Marte Vold (Master student 2011 – 2013). Management effects on low-herb diversity in outlying grasslands

Suzanne Hansen (Master student 2010 – 2012). Red-listed vascular plants species in sub-alpine and alpine landscapes: How does land-use affect their distribution?

Career and Education

2001 – 2005: BSc (Hons.) Biology, University of York

2006 – 2009: PhD Biological Sciences, University of Aberdeen

2009 - 2016: Postdoctoral Fellow & Reseacher, NTNU University Museum, Norwegian University of Science and Technology

2016 - Present: Associate Professor & Researcher, NTNU University Museum, Norwegian University of Science and Technology


Google Scholar profile

Research Gate profile


CRISTIN profile

Petersen, T.K., Vuorinen, K.E.M., Bendiksby, M. & Speed, J.D.M. (2022) Climate and land-use drive the functional composition of vascular plant assemblages across norway. Nordic Journal of Botany, e03470.

Martin, A.C., Assmann, J.J., Bradshaw, R.H.W., Kuoppamaa, M., Kuosmanen, N.I., Normand, S., Speed, J.D.M. & Macias-Fauria, M. (2022) What evidence exists for temporal variability in Arctic terrestrial and freshwater biodiversity throughout the Holocene? A systematic map protocol. Environmental Evidence, 11, 13.

Vuorinen, K., Austrheim, G., Tremblay, J.-P., Myers-Smith, I.H., Hortman, H.I., Frank, P., Barrio, I.C., Dalerum, F., Björkman, M.P., Björk, R.G., Ehrich, D., Sokolov, A., Sokolova, N., Ropars, P., Boudreau, S., Normand, S., Prendin, A.L., Schmidt, N.M., Pacheco, A., Post, E., John, C., Kerby, J.T., Sullivan, P.F., Le Moullec, M., Hansen, B.B., Van der Wal, R., Pedersen, Å.Ø., Sandal, L., Gough, L., Young, A., Li, B., Magnússon, R.Í., Sass-Klaassen, U., Buchwal, A., Welker, J.M., Grogan, P., Andruko, R., Morrissette-Boileau, C., Volkovitskiy, A., Terekhina, A. & Speed, J.D.M. (2022) Growth rings show limited evidence for ungulates’ potential to suppress shrubs across the Arctic. Environmental Research Letters.

Mienna, I.M., Austrheim, G., Klanderud, K., Bollandsås, O.M. & Speed, J.D.M. (2022) Legacy effects of herbivory on treeline dynamics along an elevational gradient. Oecologia.

Barrio, I.C., Barbero-Palacios, L., Kaarlejärvi, E., Speed, J.D.M., Heiðmarsson, S., Hik, D.S. & Soininen, E.M. (2022) What are the effects of herbivore diversity on tundra ecosystems? A systematic review protocol. Environmental Evidence, 11, 1. 10.1186/s13750-022-00257-z

Lembrechts, J.J […] Speed, J.D.M., et al. (2022), Global maps of soil temperature. Global Change Biology.

Vuorinen, K. E. M., Austrheim, G., Mysterud, A., Gya, R., Vandvik, V., Grytnes, J.-A., and Speed, J. D. M.. 2021. Functional traits of alpine plant communities show long-term resistance to changing herbivore densities. Ecosphere 12( 12):e03887. 10.1002/ecs2.3887

Kroeger, S.B., Hanslin, H.M., Lennartsson, T., D'Amico, M., Kollmann, J., Fischer, C., Albertsen, E. & Speed, J.D.M. (2021) Impacts of roads on bird species richness: A meta-analysis considering road types, habitats and feeding guilds. Science of The Total Environment, 151478.

Champagne, E., Raymond, P., Royo, A.A. Speed, J.D.M., Tremblay, J.-P., Côte, S.D. (2021). A Review of Ungulate Impacts on the Success of Climate-Adapted Forest Management Strategies. Current Forestry Reports.

Speed, J.D.M., Chimal-Ballesteros, J.A., Martin, M.D., Barrio, I.C., Vuorinen, K.E.M. & Soininen, E.M. (2021) Will borealization of Arctic tundra herbivore communities be driven by climate warming or vegetation change? Global Change Biology

Soininen, E.M., Barrio, I.C., Bjørkås, R., Björnsdóttir, K., Ehrich, D., Hopping, K.A., Kaarlejärvi, E., Kolstad, A.L., Abdulmanova, S., Björk, R.G., Bueno, C.G., Eischeid, I., Finger-Higgens, R., Forbey, J.S., Gignac, C., Gilg, O., den Herder, M., Holm, H.S., Hwang, B.C., Jepsen, J.U., Kamenova, S., Kater, I., Koltz, A.M., Kristensen, J.A., Little, C.J., Macek, P., Mathisen, K.M., Metcalfe, D.B., Mosbacher, J.B., Mörsdorf, M., Park, T., Propster, J.R., Roberts, A.J., Serrano, E., Spiegel, M.P., Tamayo, M., Tuomi, M.W., Verma, M., Vuorinen, K.E.M., Väisänen, M., van der Wal, R., Wilcots, M.E., Yoccoz, N.G. & Speed, J.D.M. (2021) Location of studies and evidence of effects of herbivory on Arctic vegetation: a systematic map. Environmental Evidence, 10, 25.2021.

Aguilar-Tomasini, M.A., Martin, M.D. and Speed, J.D.M. 2021. Assessing spatial patterns of phylogenetic diversity of Mexican mammals for biodiversity conservation. Global Ecology and Conservation 31:e01834.

Kolstad, A. L., I. B. Snøan, G. Austrheim, O. M. Bollandsås, E. J. Solberg & Speed J.D.M. 2021. Airborne laser scanning reveals increased growth and complexity of boreal forest canopies across a network of ungulate exclosures in Norway. Remote Sensing in Ecology and Conservation

Vuorinen, K.E.M., Oksanen, T., Oksanen, L., Vuorisalo, T. & Speed, J.D.M. (2021) Why don't all species overexploit? Oikos

Wana, D., Smith, S.W., Coleman, K. & Speed, J.D.M. (2021) Proximity to high densities of pastoral settlements reduces grassland regrowth in a protected tropical savanna. Biotropica

Barrio, I.C., Ehrich, D., Soininen, E.M., Ravolainen, V., Bueno, C.G., Gilg, O., Koltz, A.M., Speed, J.D.M., Hik, D. & Mörsdorf, M.A. et al. (2021) Developing common protocols to measure tundra herbivory across spatial scales. Arctic Science.

Petersen, T.K., Speed, J.D.M., Grøtan, V. & Austrheim, G. (2021) Competitors and ruderals go to town: plant community composition and function along an urbanisation gradient. Nordic Journal of Botany, 39.

Nygaard, M., Kemppainen, P., Speed, J. D.M, Elven, R., Flatberg, K. I., Galten, L. P., Yousefi, N., Solstad, H., & Bendiksby, M. Combining population genomics and ecological niche modelling to assess taxon limits between Carex jemtlandica and C. lepidocarpa. Journal of Systematics and Evolution.

Petersen, T.K., Speed, J.D.M., Grøtan, V. & Austrheim, G. (2021) Species data for understanding biodiversity dynamics: The what, where and when of species occurrence data collection. Ecological Solutions and Evidence, 2, e12048.

Tuomi, M., Väisänen, M., Ylänne, H., Brearley, F.Q., Barrio, I.C., Anne Bråthen, K., Eischeid, I., Forbes, B.C., Jónsdóttir, I.S., Kolstad, A.L., Macek, P., Petit Bon, M., Speed, J.D.M., Stark, S., Svavarsdóttir, K., Thórsson, J. & Bueno, C.G. Stomping in silence: Conceptualizing trampling effects on soils in polar tundra. Functional Ecology

Timdal, E., Westberg, M., Haugan, R., Hofton, T.H., Holien, H., Speed, J.D.M., Tønsberg, T., Bendiksby, M. (2020) Integrative taxonomy reveals a new species, Nephroma orvoi, in the N. parile species complex (lichenized Ascomycota). Graphis Scripta. vol. 32 (4).

Sundsdal A., Graae B.J., Speed J.D.M., Bukombe J., Mtweve P.J., Arneberg, M.K., Haukenes V.L., Grevskott R.T & Smith S.W. (2020) Teatime in the Serengeti: macrodetritivores sustain recalcitrant plant litter decomposition across human-modified tropical savannahs. Plant and Soil.

Vuorinen, K. E., Kolstad, A. L., De Vriendt, L., Austrheim, G., Tremblay, J. P., Solberg, E. J., & Speed, J.D.M. (2020). Cool as a moose: How can browsing counteract climate warming effects across boreal forest ecosystems? Ecology.

Speed, J.D.M., Austrheim, G., Bendiksby, M., Kolstad, A.L., & Vuorinen, K.E. (2020). Increasing Cervidae populations have variable impacts on habitat suitability for threatened forest plant and lichen species. Forest Ecology and Management, 473, 118286.

Petersen, T. K., Speed, J.D.M, Grøtan, V., & Austrheim, G. (2020). Urban aliens and threatened near-naturals: Land-cover affects the species richness of alien-and threatened species in an urban-rural setting. Scientific Reports, 10(1), 1-14.

Lembrechts, J. J., Aalto, J., Ashcroft, M. B., De Frenne, P., … Speed, J.D.M., et al. (2020). SoilTemp: a global database of near‐surface temperature. Global Change Biology.

Vuorinen, K.E.M., Rao, S.J., Hester, A.J. & Speed, J.D.M. (2020). Herbivory and climate as drivers of woody plant growth: Do deer decrease the impacts of warming? Ecological Applications.

Mienna, I.M., Speed, J.D.M., Klanderud, K., Austrheim, G., Næsset, E., & Bollandsås, O.M. (2020) The relative role of climate and herbivory in driving treeline dynamics along a latitudinal gradient. Journal of Vegetation Science. DOI: 10.1111/jvs.12865

Thomas, H.J.D., Bjorkman, A.D., Myers-Smith, I.H., ... Speed, J.D.M., et al. (2020) Global plant trait relationships extend to the climatic extremes of the tundra biome. Nature Communications, 11, 1351.

Mienna, I. M., Speed, J. D.M., Bendiksby, M., Thornhill, A. H., Mishler, B. D., & Martin, M. D. (2019). Differential patterns of floristic phylogenetic diversity across a post‐glacial landscape. Journal of Biogeography.

Halbritter, A. H., De Boeck, H. J., Eycott, A. E., Reinsch, S., Robinson, D. A., Vicca, S. ... Speed JDM et al. (2019). The handbook for standardized field and laboratory measurements in terrestrial climate change experiments and observational studies (ClimEx). Methods in Ecology and Evolution.

Smith, S.W., Graae, B.J., Bukombe, J., Hassan, S.N., Lyamuya, R.D., Jacob Mtweve, P., Treydte, A.C. & Speed, J.D.M. Savannah trees buffer herbaceous plant biomass against wild and domestic herbivores. Applied Vegetation Science, doi:10.1111/avsc.12472.

Kolstad, A.L., Austrheim, G., Graae, B.J., Solberg, E.J., Strimbeck, G.R. & Speed, J.D.M. (2019) Moose effects on soil temperatures, tree canopies, and understory vegetation: a path analysis. Ecosphere, 10, e02966.

Rheubottom, S.I., Barrio, I.C., Kozlov, M.V., ...Speed, J.D.M. ...Hik, D. S. (2019). Hiding in the background: community-level patterns in invertebrate herbivory across the tundra biome. Polar Biology,  doi:10.1007/s00300-019-02568-3

Speed, J.D.M., Austrheim, G., Kolstad, A.L. & Solberg, E.J. (2019). Long-term changes in northern large-herbivore communities reveal differential rewilding rates in space and time. PLoS ONE, 14(5), e0217166.

Speed, J.D.M., Skjelbred, I.Å., Barrio, I.C., Martin, M.D., Berteaux, D., Bueno, C.G. et al. (2019). Trophic interactions and abiotic factors drive functional and phylogenetic structure of vertebrate herbivore communities across the Arctic tundra biome. Ecography, 42,

Helsen, K., Hagenblad, J., Acharya, K.P., Brunet, J., Cousins, S.A.O., Decocq, G., De Frenne, P., Kimberley, A., Kolb, A., Michaelis, J., Plue, J., Verheyen, K., Speed, J.D.M., Graae, B.J., (2019). No genetic erosion after five generations for Impatiens glandulifera populations across the invaded range in Europe. BMC Genetics 20, 20.

Ross, L. C., Speed, J. D. M., Øien, D. I., Grygoruk, M., Hassel, K., Lyngstad, A., & Moen, A. (2019). Can mowing restore boreal rich-fen vegetation in the face of climate change?. PLOS ONE, 14(2), e0211272.

Meleshko, O., Stenøien, H. K., Speed, J. D.M., Flatberg, K. I., Kyrkjeeide, M. O., & Hassel, K. (2018). Is interspecific gene flow and speciation in peatmosses (Sphagnum) constrained by phylogenetic relationship and life-history traits?. Lindbergia, 41, linbg-01107.

Thomas, H.J.D., Myers-Smith, I.H. ..... Speed, J.D.M. ... et al. (2018) Traditional plant functional groups explain variation in economic but not size-related traits across the tundra biome. Global Ecology and Biogeography

Smith, S.W., Speed, J.D.M, Bukombe, J., Hassan, S.N., Lyamuya, R.D., Mtweve, P.J., Sundsdal, A. and Graae, B.J., 2018. Litter type and termites regulate root decomposition across contrasting savanna land‐uses. Oikos.

Bjorkman, A.D., Myers-Smith, I.H. ..... Speed, J.D.M. ... et al. (2018) Tundra Trait Team: A database of plant traits spanning the tundra biome. Global Ecology and Biogeography

Kolstad, A.L., Austrheim, G., Solberg, E. J., De Vriendt, L., & Speed, J. D.M. (2018). Pervasive moose browsing in boreal forests alters successional trajectories by severely suppressing keystone species. Ecosphere, 9(10), e02458.

Bjorkman, A.D., Myers-Smith, I.H. ..... Speed, J.D.M. ... et al. (2018) Plant functional trait change across a warming tundra biome. Nature 562, 57

Soininen, E.M., Barrio, I., Jepsen, J.U., Ehrich, D., Ravolainen, V.T. & Speed, J.D.M. (2018) Evidence of effects of herbivory on Arctic vegetation: a systematic map protocol. Environmental Evidence, 7, 23.

Thingstad, P.G., Daverdin, M. & Speed, J.D.M. (2018) Modelling Capercaillie (Tetrao urogallus) lek sites distribution and their limiting factors in a central Norwegian managed forest landscape. Ornis Norvegica, 41, 1-12.

Speed, J. D.M, Bendiksby, M., Finstad, A. G., Hassel, K., Kolstad, A. L., & Prestø, T. (2018). Contrasting spatial, temporal and environmental patterns in observation and specimen based species occurrence data. PloS one, 13(4), e0196417.

Anderson, T.M., Ngoti, P.M., Nzunda, M.L., Griffith, D.M., Speed, J.D.M, Fossøy, F., Røskaft, E. & Graae, B.J. (2018) The burning question: does fire affect habitat selection and forage preference of the black rhinoceros Diceros bicornis in East African savannahs? Oryx,

Helsen, K., Kapás, R.E., Rakvaag, G., Speed, J.D.M. & Graae, B.J. (2018) Impacts of an invasive plant on primary production: testing a functional trait-based framework with a greenhouse experiment. Journal of Vegetation Science, doi: 10.1111/jvs.12619.

Helsen, K., Smith, S.W., Brunet, J., Cousins, S.A.O., De Frenne, P., Kimberley, A., Kolb, A., Lenoir, J., Ma, S., Michaelis, J., Plue, J., Verheyen, K., Speed, J.D.M. & Graae, B.J. (2018) Impact of an invasive alien plant on litter decomposition along a latitudinal gradient. Ecosphere, 9, e02097

Kjærstad, G., Arnekleiv, J., Speed, J.D.M. & Herland, A. (2018) Effects of hydropeaking on benthic invertebrate community composition in two central Norwegian rivers. River Research and Applications.

Kolstad, A.L., Austrheim, G., Solberg, E.J., Venete, A.M.A., Woodin, S.J. & Speed, J.D.M. (2017) Cervid Exclusion Alters Boreal Forest Properties with Little Cascading Impacts on Soils. Ecosystems 10.1007/s10021-017-0202-4.

Graae, B.J., Vandvik, V., Armbruster, W.S., Eiserhardt, W.L., Svenning, J.-C., Hylander, K., Ehrlén, J., Speed, J.D.M., Klanderud, K., Bråthen, K.A., Milbau, A., Opedal, Ø.H., Alsos, I.G., Ejrnæs, R., Bruun, H.H., Birks, H.J.B., Westergaard, K.B., Birks, H.H. & Lenoir, J. Stay or go – how topographic complexity influences alpine plant population and community responses to climate change. Perspectives in Plant Ecology, Evolution and Systematics

Barrio, I.C., Lindén, E., Te Beest, M., Olofsson, J., Rocha, A., Soininen, E.M., Alatalo, J.M., Andersson, T., Asmus, A., Boike, J., Bråthen, K.A., Bryant, J.P., Buchwal, A., Bueno, C.G., Christie, K.S., Denisova, Y.V., Egelkraut, D., Ehrich, D., Fishback, L., Forbes, B.C., Gartzia, M., Grogan, P., Hallinger, M., Heijmans, M.M.P.D., Hik, D.S., Hofgaard, A., Holmgren, M., Høye, T.T., Huebner, D.C., Jónsdóttir, I.S., Kaarlejärvi, E., Kumpula, T., Lange, C.Y.M.J.G., Lange, J., Lévesque, E., Limpens, J., Macias-Fauria, M., Myers-Smith, I., van Nieukerken, E.J., Normand, S., Post, E.S., Schmidt, N.M., Sitters, J., Skoracka, A., Sokolov, A., Sokolova, N., Speed, J.D.M., Street, L.E., Sundqvist, M.K., Suominen, O., Tananaev, N., Tremblay, J.-P., Urbanowicz, C., Uvarov, S.A., Watts, D., Wilmking, M., Wookey, P.A., Zimmermann, H.H., Zverev, V. & Kozlov, M.V. (2017) Background invertebrate herbivory on dwarf birch (Betula glandulosa-nana complex) increases with temperature and precipitation across the tundra biome. Polar Biology, DOI 10.1007/s00300-017-2139-7.

Wangen, K., Speed, J.D.M. & Hassel, K. (2017) How does microclimate affect the growth of the rare liverwort Scapania nimbosa? Lindbergia, 40, 1-10.

Speed, J.D.M., & Austrheim, G. (2017). The importance of herbivore density and management as determinants of the distribution of rare plant species. Biological Conservation, 205, 77-84.

Anderson, H.B., Speed, J.D.M., Madsen, J., Pedersen, Å.Ø., Tombre, I.M., & Van der Wal, R. (2016). Late snow melt moderates herbivore disturbance of the Arctic tundra. Ecoscience, 23(1-2), 29-39.

Austrheim, G., Speed, J.D.M., Evju, M., Hester, A., Holand, Ø., Loe, L.E., Martinsen, V., Mobæk, R., Mulder, J., Steen, H., 2016. Synergies and trade-offs between ecosystem services in an alpine ecosystem grazed by sheep–an experimental approach. Basic and Applied Ecology 10.1016/j.baae.2016.06.003.

Barrio, I.C., Bueno, C.G., Gartzia, M., Soininen, E.M., Christie, K.S., Speed, J.D.M., Ravolainen, V.T., Forbes, B.C., Gauthier, G., Horstkotte, T., Hoset, K.S., Høye, T.T., Jónsdóttir, I.S., Lévesque, E., Mörsdorf, M.A., Olofsson, J., Wookey, P.A. & Hik, D.S. (2016). Biotic interactions mediate patterns of herbivore diversity in the Arctic. Global Ecology and Biogeography, 10.1111/geb.12470.

Wangen, K., Speed, J.D.M., & Hassel, K. (2016). Hyper-oceanic liverwort species of conservation concern: evidence for dispersal limitation and identification of suitable uncolonised regions. Biodiversity and Conservation, 25: 1053-1071

Ross, L.C., Austrheim, G., Asheim, L.-J., Bjarnason, G., Feilberg, J., Fosaa, A.M., Hester, A.J., Holand, Ø., Jónsdóttir, I.S., Mortensen, L.E., Mysterud, A., Olsen, E., Skonhoft, A., Speed, J.D.M., Steinheim, G., Thompson, D.B.A. & Thórhallsdóttir, A.G. (2016). Sheep grazing in the North Atlantic region: A long-term perspective on environmental sustainability. Ambio, 1-16.

Wasof, S., Lenoir, J., Aarrestad, P.A., Alsos, I.G., Armbruster, W.S., Austrheim, G., Bakkestuen, V., Birks, H.J.B., Bråthen, K.A., Broennimann, O., Brunet, J., Bruun, H.H., Dahlberg, C.J., Diekmann, M., Dullinger, S., Dynesius, M., Ejrnæs, R., Gégout, J.-C., Graae, B.J., Grytnes, J.-A., Guisan, A., Hylander, K., Jónsdóttir, I.S., Kapfer, J., Klanderud, K., Luoto, M., Milbau, A., Moora, M., Nygaard, B., Odland, A., Pauli, H., Ravolainen, V., Reinhardt, S., Sandvik, S.M., Schei, F.H., Speed, J.D.M., Svenning, J.-C., Thuiller, W., Tveraabak, L.U., Vandvik, V., Velle, L.G., Virtanen, R., Vittoz, P., Willner, W., Wohlgemuth, T., Zimmermann, N.E., Zobel, M. & Decocq, G. (2015) Disjunct populations of European vascular plant species keep the same climatic niches. Global Ecology and Biogeography, 10.1111/geb.12375.

Myers-Smith, I.H., Elmendorf, S., Beck, P., Wilmking, M., Hallinger, M., Blok, D., Tape, K., Rayback, S.A., Macias-Fauria, M., Forbes, B.C., Speed, J.D.M., Boulanger-Lapointe, N., Rixen, C., Levesque, E., Schmidt, N., Baittinger, C., Trant, A., Hermanutz, L., Siegwart Collier, L., Dawes, M., Lantz, T., Weijers, S., Jørgensen, R., Buchwal, A., Buras, A., Naito, A.T., Ravolainen, V., Schaepman-Strub, G., Wheeler, J., Wipf, S., Guay, K., Hik, D.S. & Velland, M. (2015) Climate sensitivity of shrub growth across the tundra biome. Nature Climate Change, 5: 887-891.

Anderson, H.B., Hubner, C.E., Speed, J.D.M., Madsen, J. & Van Der Wal, R. (2015) Biding time before  breeding: flexible use of the Arctic landscape by migratory geese during spring. Polar Research, 34, 26372.

Bernes, C., Carpenter, S.R., Gårdmark, A., Larsson, P., Persson, L., Skov, C., Speed, J.D.M. & Van Donk, E. (2015) What is the influence of a reduction of planktivorous and benthivorous fish on water quality in temperate eutrophic lakes? A systematic review. Environmental Evidence, 4, 7.

Speed, J. D. M., Martinsen, V., Hester, A. J., Holand, Ø., Mulder, J., Mysterud, A., and Austrheim, G. (2015) Continuous and discontinuous variation in ecosystem carbon stocks with elevation across a treeline ecotone, Biogeosciences, 12, 1615-1627.

Bernes, C., Bråthen, K.A., Forbes, B.C., Speed, J.D.M. & Moen, J. (2015) What are the impacts of reindeer/caribou (Rangifer tarandus L.) on arctic and alpine vegetation? Environmental Evidence, 4, 4.

Kjærstad, G., Arnekleiv, J.V. & Speed, J.D.M. (2015) Effects of Three Consecutive Rotenone Treatments on the Benthic Macroinvertebrate Fauna of the River Ogna, Central Norway. River Research and Applications, 10.1002/rra.2873.

Thingstad, P.G., Hogstad, O. & Speed, J.D.M. (2014) The influence of climatic conditions in breeding grounds and migratory flyways on a subalpine Norwegian Willow Warbler Phylloscopus trochilus population. Ornis Fennica

Austrheim G., Speed J.D.M., Martinsen V., Mulder J., Mysterud A. (2014). Experimental Effects of Herbivore Density on Aboveground Plant Biomass in an Alpine Grassland Ecosystem. Arctic, Antarctic, and Alpine Research 46: 535-541.

Speed J.D.M., Martinsen V., Mysterud A., Mulder J., Holand Ø., Austrheim G. (2014). Long-Term Increase in Aboveground Carbon Stocks Following Exclusion of Grazers and Forest Establishment in an Alpine Ecosystem. Ecosystems 10.1007/s10021-014-9784-2.

Speed, J.D.M., Austrheim, G., Hester, A.J., Meisingset, E., Mysterud, A, Tremblay J.-P. Øien D.-I. & Solberg, E.J. (2014). General and specific responses of understory vegetation to cervid herbivory across a range of boreal forests. Oikos. DOI: 10.1111/oik.01373

Sjögren, P., Solem, T., Stenvik, L., Tretvik, A.M., Daverdin, M., Speed, J.D.M., Austrheim, G. (2014) Quantification of interpreted pollen data based on the R-value model – exemplified by impact assessment of pre-historic iron production and summer farming in Budalen, central Norway. The Holocence. 10.1177/0959683613520256

Miljeteig, C., Olsen, A.J., Båtnes, A.S., Altin, D., Nordtug, T., Alver, M.O., Speed, J.D.M. & Jenssen, B.M. (2014) Sex and life stage dependent phototactic response of the marine copepod Calanus finmarchicus (Copepoda: Calanoida). Journal of Experimental Marine Biology and Ecology, 451, 16-24.

Speed, J.D.M., Meisingset, E., Austrheim, G., Hester, A.J., Mysterud, A, Tremblay J.-P. & Solberg, E.J. (2013). Low intensities of red deer browsing constrain rowan growth in mature boreal forests of western Norway. Ecoscience. 20:3. 311-318.

Pedersen, Å. Ø., Speed, J. D. M., and Tombre, I. M., (2013). Prevalence of grubbing in the arctic tundra increases with the pink-footed goose population expansion. Polar Biology, 10.1007/s00300-013-1374-9.

Speed, J.D.M., Austrheim, G. & Mysterud, A. (2013) The response of plant diversity to grazing varies along an elevational gradient. Journal of Ecology, 101: 1225-1236.

Speed, J.D.M., Austrheim, G., Hester, A.J. & Mysterud, A. (2013) The response of alpine Salix shrubs to long-term browsing varies with elevation and herbivore density. Arctic Antarctic and Alpine Research. 45: 584-593.

Bernes, C., Bråthen, K.A., Forbes, B.C., Hofgaard, A., Moen, J. & Speed, J.D.M. (2013). What are the impacts of reindeer/caribou (Rangifer tarandus L.) on arctic and alpine vegetation? A systematic review protocol. Environmental Evidence, 2, 6.

Lenoir, J., Graae, B.J., Aarrestad, P.A., Alsos, I.G., Armbruster, S.W., Austrheim, G., Bergendorff, C., Birks, H.J.B., Bråthen, K.A., Brunet, J., Bruun, H.H., Dahlberg, C.J., Decocq, G., Diekmann, M., Dynesius, M., Ejrnæs, R., Grytnes, J.-A., Hylander, K., Klanderud, K., Luoto, M., Milbau, A., Moora, M., Nygaard, B., Odland, A., Ravolainen, V.T., Reinhardt, S., Sandvik, S.M., Schei, F.H., Speed, J.D.M., Tveraabak, L.U., Vandvik, V., Velle, L.G., Virtanen, R., Zobel, M., Svenning, J.-C., 2013. Local temperatures inferred from plant communities suggest strong spatial buffering of climate warming across Northern Europe. Global Change Biology, 10.1111/gcb.12129.

Myking, T., Solberg, E.J., Austrheim, G., Speed, J.D.M., Bøhler, F., Astrup, R. & Eriksen, R. (2013) Browsing of sallow (Salix caprea L.) and rowan (Sorbus aucuparia L.) in the context of life history strategies: a literature review. European Journal of Forest Research, DOI 10.1007/s10342-013-0684-3.

Speed, J.D.M., Austrheim, G., Hester, A.J., Solberg, E.J. & Tremblay, J.-P. (2013) Regional-scale alteration of clear-cut forest regeneration caused by moose browsing. Forest Ecology and Management, 289, 289-299.

Speed, J.D.M., Austrheim, G., Birks, H.J.B., Johnson, S., Kvamme, M., Nagy, L., Sjögren, P., Skar, B., Stone, D., Svensson, E. & Thompson, D.B.A. (2012) Natural and cultural heritage in mountain landscapes: towards an integrated valuation. International Journal of Biodiversity Science, Ecosystem Services & Management, 8, 313-320.

Speed, J.D.M., Austrheim, G., Hester, A.J. & Mysterud, A. (2012) Elevational advance of alpine plant communities is buffered by herbivory. Journal of Vegetation Science 23:617-625

Speed, J.D.M., Austrheim, G., Hester, A.J. & Mysterud, A. (2011) Browsing interacts with climate to determine tree-ring increment. Functional Ecology. 25, 1018-1023

Speed, J.D.M., Austrheim, G., Hester, A.J. & Mysterud, A. (2011) Growth limitation of mountain birch caused by sheep browsing at the altitudinal treeline. Forest Ecology and Management, 261, 1344-1352.

Speed, J.D.M., Cooper, E.J., Jónsdóttir, I.S., Van der Wal, R. & Woodin, S.J. (2010) Plant community properties predict vegetation resilience to herbivore disturbance in the Arctic. Journal of Ecology. 98, 1002-1013. DOI 10.1111/j.1365-2745.2010.01685.x

Speed, J.D.M., Austrheim, G., Hester, A.J. & Mysterud, A. (2010) Experimental evidence for herbivore limitation of the treeline. Ecology 91, 3414-3420. DOI 10.1890/09-2300

Speed, J.D.M., Woodin, S.J., Tømmervik, H. & Van der Wal, R. (2010) Extrapolating herbivore-induced carbon loss across an arctic landscape. Polar Biology, 33, 789-797 DOI 10.1007/s00300-009-0756-5

Speed, J.D.M., Woodin, S.J., Tommervik, H., Tamstorf, M.P. & Van der Wal, R. (2009) Predicting Habitat Utilization and Extent of Ecosystem Disturbance by an Increasing Herbivore Population. Ecosystems, 12, 349-359.

Speed, J.D.M., Van der Wal, R. & Woodin, S.J. (2009) Do Nitrogen Concentration and Forage Quality of the Moss Racomitrium lanuginosum Increase with Latitude? Research Letters in Ecology, vol. 2009, 4.

Scientific, academic and artistic work

A selection of recent journal publications, artistic productions, books, including book and report excerpts. See all publications in the database

Journal publications